蒂奥得螺属
腹足綱蜑螺科軟體動物的一個屬
蒂奥得螺属（学名：Theodoxus）是腹足纲蜑螺科软体动物的一个属，体型细小，有螺厣，栖息于淡水或半咸水环境中[2]。
蒂奥得螺属 化石时期：
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| An oblique left side view of a live Theodoxus fluviatilis | |
科学分类 
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| 界： | 动物界 Animalia |
| 门： | 软体动物门 Mollusca |
| 纲： | 腹足纲 Gastropoda |
| 目： | 蜑形目 Cycloneritida |
| 总科： | 蜑螺总科 Neritoidea |
| 科： | 蜑螺科 Neritidae |
| 亚科： | 蜑螺亚科 Neritinae |
| 属： | 蒂奥得螺属 Theodoxus Montfort, 1810[1] |
| 模式种 | |
| Theodoxus lutetianus Montfort, 1810 | |
| 异名[2] | |
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分布
本属物种分布于欧洲及北非，并往东延伸至伊朗南部[3]。除本属以外，蜑螺科并没有其他属与本属具同域分布[4]。这在蜑螺科相当例外，因为其他蜑螺科物种只分布于南半球的热带[4]。本属物种均为蜑螺科仅有栖息于温带的物种[4]。
Bunje 和 Lindberg (2007) 提出了Theodoxus分支的第一个系统发育假说。 [4] Theodoxus属的进化受到中新世地中海与副特提斯海分离的影响。 [4]
形态描述
本属物种的螺壳呈半卵形，具有平坦的壳口。没有螺脐。螺柱及内螺层溶解[3]。
蒂奥得螺属物种间的差异，无论是大小、外壳膜上的颜色与花纹变化、螺厣及齿舌的细节都有很大差异，这使得本属物种的辨认比较困难[5]。
生态
These animals live on stones, and often also under stones, in up to 5–6 m depth or deeper, feeding on algal covers.[3] Theodoxus needs rough surfaces in order to be able to digest its food, so a stony 基质 (生物生活环境) is necessary.[3] Green algae [需要解释] are not consumed; Theodoxus has no 纤维素酶s.[3] These snails lay egg capsules containing 30-70 eggs each, usually on the shells of other Theodoxus animals; only one juvenile grows, the other eggs serve as food.[3]
物种
截至2007年，还未有对蒂奥得螺属作系统评价的研究[4]。Bunje (2004) 的研究指出本属至少有34个现害种；而在2015年度的国际自然保护联盟濒危物种红色名录列出了本属包括有至少23个物种[6]。
本属包括下列各物种：
- Subgenus Brusinaella Andrusov, 1912[7]
- † Theodoxus petasatus (Seninski, 1905)
- Subgenus Calvertia Bourguignat, 1880[8]
- † Theodoxus acuticarinatus (Fuchs, 1870)
- † Theodoxus amblygonioides Wenz, 1930
- † Theodoxus amethystinus (Brusina, 1874)
- † Theodoxus anconae (Capellini, 1880)
- † Theodoxus angularis Gozhik, 2002
- † Theodoxus anonymus Anistratenko & Gozhik, 1995
- † Theodoxus atticus Kühn, 1963
- † Theodoxus barakovici (Brusina, 1902)
- † Theodoxus bessarabicus (Sinzov, 1896)
- † Theodoxus brusinai (Pavlović, 1931)
- † Theodoxus capellinii (Pantanelli, 1876)
- † Theodoxus capillaceus (Brusina, 1874)
- † Theodoxus crescens (Fuchs, 1870)
- † Theodoxus doderleini (d'Ancona, 1869)
- † Theodoxus dumortieri (Fontannes, 1878)
- † Theodoxus fasciatus Gozhik in Gozhik & Datsenko, 2007
- † Theodoxus gnezdai (Brusina, 1884)
- † Theodoxus grateloupianus (Férussac, 1823)
- † Theodoxus hoernesanus (Semper, 1867)
- † Theodoxus ingulenzis Gozhik in Gozhik & Datsenko, 2007
- † Theodoxus kalodictya (Andrusov, 1909)
- † Theodoxus koslinskyi (Porumbaru, 1881)
- † Theodoxus lamellatus (Brusina, 1892)
- † Theodoxus lineatus (Sinzov, 1896)
- † Theodoxus lorkovici (Brusina, 1878)
- † Theodoxus macedonicus Wenz, 1943
- † Theodoxus maculosus Gozhik in Gozhik & Datsenko, 2007
- † Theodoxus mayeri (Semper, 1867)
- † Theodoxus militaris (Neumayr, 1869)
- † Theodoxus miljkovici (Brusina, 1902)
- † Theodoxus millepunctatus (Brusina, 1902)
- † Theodoxus moeschi Locard, 1893
- † Theodoxus morulus Kühn, 1963
- † Theodoxus mutinensis (d'Ancona, 1869)
- † Theodoxus neumayri (Burgerstein, 1877)
- † Theodoxus nitens Gozhik, 2002
- † Theodoxus nivosus (Brusina, 1874)
- † Theodoxus novorossicus (Sinzov, 1896)
- † Theodoxus obtusangulaeformis Gozhik, 2002
- † Theodoxus oxytropida (Andrusov, 1909)
- † Theodoxus pavlovici Milošević, 1983
- † Theodoxus philippianus Locard, 1883
- † Theodoxus pilari (Brusina, 1884)
- † Theodoxus platystoma (Brusina, 1874)
- † Theodoxus pseudodanubialis (Sinzov, 1896)
- † Theodoxus pseudofluviatilis Locard, 1893
- † Theodoxus pseudograteloupanus (Sinzov, 1884)
- † Theodoxus radmanesti (Fuchs, 1870)
- † Theodoxus radovanovici (Brusina, 1893)
- † Theodoxus reiseri (Brusina, 1902)
- † Theodoxus rugosus (Pavlović, 1931)
- † Theodoxus sagittiferus (Brusina, 1874)
- † Theodoxus schachmaticus (Andrusov, 1909)
- † Theodoxus scoliogramma (Brusina, 1884)
- † Theodoxus semidentatus (Sandberger, 1875)
- † Theodoxus serrulatus (Brusina, 1892)
- † Theodoxus slavonicus (Brusina, 1878)
- † Theodoxus sophievkaensis Gozhik in Gozhik & Datsenko, 2007
- † Theodoxus stanae (Brusina, 1893)
- † Theodoxus stefanescui (Fontannes, 1887)
- † Theodoxus stefanescuiformis Gozhik in Gozhik & Datsenko, 2007
- † Theodoxus subquadrofasciatus Gozhik, 2002
- † Theodoxus subslavonicus Gozhik, 2002
- † Theodoxus sundicus (Andrusov, 1909)
- † Theodoxus suskalovici (Pavlović, 1903)
- † Theodoxus sycophantus (Brusina, 1878)
- † Theodoxus tropidophorus (Brusina, 1884)
- † Theodoxus turbinatus (Fuchs, 1870)
- † Theodoxus veljetinensis (Pavlović, 1903)
- † Theodoxus venustus (Brusina, 1897)
- † Theodoxus xanthozona (Brusina, 1884)
- † Theodoxus zagradovkaensis Gozhik, 2002
- † Theodoxus zivkovici (Pavlović, 1903)
- Subgenus Neritonyx Andrusov, 1912[9]
- † Theodoxus unguiculatus (Seninski, 1905)
- Subgenus Ninniopsis Tomlin, 1930[10]
- † Theodoxus colchicus (Andrusov, 1912)
- Subgenus Neritaea Roth, 1855[11]
- Theodoxus anatolicus (Récluz, 1841)[12][13][14]
- † Theodoxus groyanus (Férussac, 1823)
- Theodoxus jordani (Sowerby)[14]
- † Theodoxus micans (Gaudry & Fischer in Gaudry, 1867)
- Theodoxus niloticus (Reeve, 1856)[14] - synonym: Theodoxus africanus (Reeve, 1856)[15]
- † Theodoxus subdoricus Schütt, 1976
- Theodoxus subterrelictus Schütt, 1963[12][13]
- Theodoxus peloponensis (Récluz, 1841)
- Theodoxus varius (Menke, 1828)[12][13]
- Subgenus Theodoxus Montfort, 1810
- Theodoxus altenai Schütt, 1965[12][14]
- Theodoxus baeticus (Lamarck, 1822)[12][16]
- † Theodoxus becenensis (Cobălcescu, 1883)
- † Theodoxus brenneri (Handmann, 1882)
- † Theodoxus carasiensis Jekelius, 1944
- † Theodoxus constantiae Stefanescu, 1896
- † Theodoxus culceri (Porumbaru, 1881)
- † Theodoxus cyrtocelis (Krauss, 1852)
- Theodoxus danubialis (Pfeiffer, 1828)[12][16][14]
- † Theodoxus denisluensis (Oppenheim, 1919)
- † Theodoxus deperditus (Almera, 1894)
- † Theodoxus elongatulus (Philippi, 1844) [12][16]
- † Theodoxus eugenii Jekelius, 1944
- Theodoxus euxinus (Clessin, 1886)[12][16][17]
- Theodoxus fluviatilis (Linnaeus, 1758)[12][16] - 模式种.[18] synonyms: Theodoxus brauneri (Lindholm, 1908)[18] (but as separate species per[12][16]); Theodoxus brauneri f. lacrymans Lindholm, 1908;[18] Theodoxus brauneri f. alboguttata Lindholm, 1908;[18] Theodoxus brauneri f. pulherrima Lindholm, 1908;[18] Theodoxus lutetianus Montfort, 1810.[3] Bunje (2005)[19] does not consider Theodoxus velox Anistratenko, 1999[20][17] to be a distinct species from Theodoxus fluviatilis.[19][14][17]
- † Theodoxus gregarius (Thomä, 1845)
- Theodoxus heldreichi (Martens, 1879)[12][14]
- † Theodoxus hisingeri (Bellardi & Michelotti, 1841)
- Theodoxus hispalensis (Martens, 1879)[12][16]
- † Theodoxus jekeliusi Jurišić-Polšak, 1979
- † Theodoxus leobersdorfensis (Handmann, 1887)
- Theodoxus meridionalis (Philippi, 1836)[12][21][16]
- † Theodoxus milessii Papp, 1979
- † Theodoxus moosbrunnensis Papp, 1953
- † Theodoxus morellii (Bellardi & Michelotti, 1841)
- Theodoxus pallasi Lindholm, 1924[22][12][16][17] - synonym: Theodoxus lituratus (Eichwald, 1838)[22]
- † Theodoxus pilidei (Tournouër, 1879)
- Theodoxus prevostianus (Pfeiffer, 1828)[12][16][14][23]
- † Theodoxus punctatolineatus (Sinzov, 1896)
- Theodoxus saulcyi (Bourguignat, 1852)[12][16]
- † Theodoxus scamandrius (Calvert & Neumayr, 1880)
- † Theodoxus semiplicatus (Neumayr in Herbich & Neumayr, 1875)
- † Theodoxus simplex (Fuchs, 1877)
- † Theodoxus subglobosus (Eichwald, 1853)
- Theodoxus subthermalis (Bourguignat in Issel, 1865)[17] or Theodoxus fluviatilis subthermalis Issel, 1865[16]
- Theodoxus transversalis (Pfeiffer, 1828)[12][16][14]
- † Theodoxus trifasciatus (Grateloup, 1839)
- † Theodoxus turislavicus Jekelius, 1944
- Theodoxus valentinus (Graells, 1846)[12][16]
- † Theodoxus zografi (Brusina, 1902)
- Subgenus ?
- † Theodoxus abnormis (Jenkins, 1864)
- † Theodoxus almelae (Revilla, 1958)
- Theodoxus astrachanicus Starobogatov, 1994[22][17]
- † Theodoxus banaticus Jekelius, 1944
- † Theodoxus bohotinensis (Simionescu & Barbu, 1940)
- † Theodoxus bolivari (Royo Gómez, 1922)
- † Theodoxus bukowskii (Oppenheim, 1919)
- Theodoxus cariosus (Wood, 1828)
- Theodoxus coronatus (Leach, 1815)[24]
- † Theodoxus crenulatus (Klein, 1853)
- † Theodoxus cunici (Brusina, 1892)
- Theodoxus danasteri (Lindholm, 1908)[20][17]
- † Theodoxus doricus (Neumayr, 1880)
- Theodoxus euxinus (Clessin, 1886)
- Theodoxus gloeri Odabaşi & Arslan, 2015
- Theodoxus gurur Sands & Glöer, 2020
- † Theodoxus hellenicus (Bukowski, 1896)
- † Theodoxus imbricatus (Brusina, 1878)
- † Theodoxus intracarpaticus Jekelius, 1944
- Theodoxus karasuna[来源请求]
- † Theodoxus licherdopoli (Stefanescu, 1896)
- Theodoxus luteofasciatus (Miller, 1879)
- Theodoxus macri (Sowerby, 1849)[来源请求]
- Theodoxus maresi (Bourguignat, 1864)[25]
- Theodoxus marteli (Pallary, 1920)[26]
- Theodoxus milachevitchi Golikov & Starobogatov, 1966[17]
- Theodoxus mutinensis[来源请求]
- Theodoxus numidicus (Récluz, 1841)[27]
- Theodoxus olivaceus[来源请求]
- Theodoxus pallasi Lindholm, 1924
- Theodoxus pallidus Dunker, 1861
- † Theodoxus pappi Sauerzopf, 1952
- † Theodoxus paradisii (Magrograssi, 1928)
- † Theodoxus patrae Esu & Girotti, 2015
- † Theodoxus percarinatus (Oppenheim, 1919)
- Theodoxus pilidei (Tournouêr, 1879)[12]
- † Theodoxus politioanei Jekelius, 1944
- † Theodoxus politus Jekelius, 1944
- Theodoxus poppei Ahuir, 2020
- † Theodoxus postcrenulatus Papp, 1953
- † Theodoxus prozlatarici Jekelius, 1944
- † Theodoxus pseudodacicus Neubauer, Harzhauser, Georgopoulou, Mandic & Kroh, 2014 †
- † Theodoxus quadrifasciatus (Bielz, 1864) †
Theodoxus reticularis Sowerby, 1838[28] IT IS A SYNONYM OF Clithon reticularis http://oldredlist.iucnredlist.org/details/166739/0[失效链接] -->
- † Theodoxus reticulatus Pană, 1990
Theodoxus retropictus UNREFERENCED - see probably Clithon retropictum -->
- † Theodoxus rhodiensis (Tournouër in Fischer, 1877)
- Theodoxus sarmaticus (Lindholm, 1901)[20][17]
- Theodoxus schultzii (Grimm, 1877)[22][17]
- † Theodoxus scriptus (Stefanescu, 1896)
- † Theodoxus serratiliniformis Geyer, 1914[23]
- † Theodoxus sinjanus (Brusina, 1876)
- † Theodoxus soceni Jekelius, 1944
- † Theodoxus sphaeroidalis (Revilla, 1958)
- † Theodoxus spratti (Jenkins, 1864)
- † Theodoxus stoicai Neubauer, Harzhauser, Georgopoulou, Mandic & Kroh, 2014
- Theodoxus subthermalis Bourguignat in Issel, 1865
- † Theodoxus susuzianus Harzhauser, Neubauer & Hoşgör, 2018
- † Theodoxus timisensis Jekelius, 1944
- † Theodoxus trilophosensis Rust, 1997
- Theodoxus velascoi (Graëlls, 1846)[12]
- Theodoxus velox V. Anistratenko in O. Anistratenko, Starobogatov & V. Anistratenko, 1999
- Theodoxus vespertinus (Sowerby II, 1849)
- Theodoxus wesselinghi Sands & Glöer, 2020
- Theodoxus wilkei Sands & Glöer, 2020
- † Theodoxus zlatarici Brusina, 1902)
- Synonyms
- † Theodoxus burdigalensis (d'Orbigny, 1852) : synonym of † Theodoxus (Theodoxus) trifasciatus (Grateloup, 1839)
- Theodoxus cariosus (Wood, 1828) and Theodoxus cariosa are synonyms of Neritina cariosa (Wood, 1828)[29]
- † Theodoxus dacicus Jekelius, 1944: synonym of † Theodoxus (Theodoxus) leobersdorfensis dacicus Jekelius, 1944
- Theodoxus doriae Issel, 1865: synonym of Theodoxus fluviatilis (Linnaeus, 1758) represented as Theodoxus (Theodoxus) fluviatilis (Linnaeus, 1758)
- Theodoxus euphraticus (Mousson, 1874)[30] is a synonym of Neritina euphratica Mousson, 1874[31]
- † Theodoxus intracarpatica Jekelius, 1944 : synonym of † Theodoxus intracarpaticus Jekelius, 1944
- Theodoxus lutetianus Montfort, 1810: synonym of Theodoxus fluviatilis (Linnaeus, 1758) represented as Theodoxus (Theodoxus) fluviatilis (Linnaeus, 1758)
- † Theodoxus mariae (Handmann, 1887): synonym of † Neritina mariae Handmann, 1887
- † Theodoxus martensi (Brusina, 1884): synonym of † Ninnia martensi (Brusina, 1884)
- Theodoxus neglectus is a synonym of Neritina neglecta (Pease, 1861)[29]
- † Theodoxus oslavanensis Rzehak, 1893: synonym of † Theodoxus (Theodoxus) cyrtocelis austriacus (Rzehak, 1893)
- Theodoxus oualaniense (Lesson, 1831): synonym of Checked: verified by a taxonomic editorClithon oualaniense (Lesson, 1831)
- † Theodoxus perlongus Jekelius, 1944: synonym of † Theodoxus (Theodoxus) zografi perlongus Jekelius, 1944
- † Theodoxus petralbensis Jekelius, 1944: synonym of † Theodoxus (Theodoxus) zografi petralbensis Jekelius, 1944
- † Theodoxus pictus (Férussac, 1823): synonym of † Agapilia picta (Férussac, 1823)
- Theodoxus reclivatus Say, 1822: synonym of Neritina usnea (Röding, 1798)
- † Theodoxus rumanus (Stefanescu, 1896): synonym of † Theodoxus (Calvertia) stefanescui (Fontannes, 1887)
- † Theodoxus scamandri (Calvert & Neumayr, 1880): synonym of † Theodoxus (Theodoxus) scamandrius (Calvert & Neumayr, 1880)
- Theodoxus vespertinus : synonym of Neritina vespertinus (Sowerby, 1849)[29]
- † Theodoxus vetranici (Brusina, 1902): synonym of † Neritina vetranici (Brusina, 1902)
- Theodoxus virginea (Linnaeus, 1758): synonym of Neritina virginea (Linnaeus, 1758)
- † Theodoxus (Vittoclithon) pictus bizonalis (Grateloup, 1839): synonym of † Agapilia picta bizonalis (Grateloup, 1839)
参考文献
- ^ Montfort D. de (1810). Conchyliologie systématique, et classification méthodique des coquilles; offrant leurs figures, leur arrangement générique, leurs descriptions caractéristiques, leurs noms; ainsi que leur synonymie en plusieurs langues. Ouvrage destiné à faciliter l'étude des coquilles, ainsi que leur disposition dans les cabinets d'histoire naturelle. Coquilles univalves, non cloisonnées. Tome second. pp. [1-3], 1-676. Paris. (Schoell). page 350.
- ^ 2.0 2.1 Bouchet, P. (2014). Theodoxus Montfort, 1810. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=153531 （页面存档备份，存于互联网档案馆） on 2014-11-17.
- ^ 3.0 3.1 3.2 3.3 3.4 3.5 3.6 3.7 "Genus summary for Theodoxus " （页面存档备份，存于互联网档案馆）. AnimalBase, last modified 24 February 2009, accessed 11 April 2011.
- ^ 4.0 4.1 4.2 4.3 4.4 4.5 Bunje P. M. & Lindberg D. R. (2007). "Lineage divergence of a freshwater snail clade associated with post-Tethys marine basin development". Molecular Phylogenetics and Evolution 42(2): 373–387. doi:10.1016/j.ympev.2006.06.026doi:10.1016/j.ympev.2006.06.026.
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- ^ The IUCN Red List of Threatened Species. Version 2014.3. <www.iucnredlist.org>. Downloaded on 23 March 2015. (Searching for "Theodoxus".)
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- ^ Andrussov (1912). Verh. Russ. min. Ges. St. Pétersburg (2)49: page 4 (Brusinaella) and page 1 (Neritonyx).
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- ^ Roth (1855). Malak. Bl. 2: 56.
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- ^ Van Damme, D.; Ghamizi, M. Theodoxus maresi. The IUCN Red List of Threatened Species. 2010, 2010: e.T21727A9314600 [17 November 2021]. doi:10.2305/IUCN.UK.2010-3.RLTS.T21727A9314600.en .
- ^ Van Damme, D.; Ghamizi, M. Theodoxus marteli. The IUCN Red List of Threatened Species. 2010, 2010: e.T184623A8302935 [17 November 2021]. doi:10.2305/IUCN.UK.2010-3.RLTS.T184623A8302935.en .
- ^ Van Damme, D.; Ghamizi, M. Theodoxus numidicus. The IUCN Red List of Threatened Species. 2010, 2010: e.T21728A9314774 [17 November 2021]. doi:10.2305/IUCN.UK.2010-3.RLTS.T21728A9314774.en .
- ^ Madhyastha A., Dutta J. & Daniel B. A. (2010). Theodoxus reticularis[失效链接]. In: IUCN 2010. IUCN Red List of Threatened Species. Version 2010.4. <www.iucnredlist.org>. Downloaded on 11 April 2011.
- ^ 29.0 29.1 29.2 Goodwin D. R. (2006). "The Discovery of Neritina (Theodoxus ) cariosa (Wood, 1828) on the Island of Maui, Hawaii (Gastropoda: Neritidae)". Visaya Net 11 pp. PDF （页面存档备份，存于互联网档案馆）.
- ^ Mansoorian A. (2001). "Freshwater Gastropod of Khuzestan Province, South-West Iran". Iranian Int. J. Sci. 2(2): 9 pp. PDF （页面存档备份，存于互联网档案馆）.
- ^ Glöer P. & Pešić V. (2012). "The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species". ZooKeys 219: 11-61, doi:10.3897/zookeys.219.3406.
- Eichhorst T.E. (2016). Neritidae of the world. Volume 2. Harxheim: Conchbooks. Pp. 696-1366
外部链接
- Bandel, K. The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda). Mittelungen aus dem Geologisch-Palaontologischen Institut Universitat Hamburg. 2001, 85: 65–164 [2022-10-21]. 原始内容存档于2011-07-26 （英语）.