剑水蚤属
剑水蚤科的一属甲壳动物
剑水蚤属（学名：Cyclops）是剑水蚤科下的一个属，其物种亦是最常见的淡水桡脚类生物，包括有超过400个不同物种[1][2]，当中在大中华地区有发现的有三种。 连同其他体型同样微小的非桡脚类甲壳亚门物种，特别是枝角目，常被泛称为“水蚤”。
剑水蚤属
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科学分类 
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| 界： | 动物界 Animalia |
| 门： | 节肢动物门 Arthropoda |
| 亚门： | 甲壳亚门 Crustacea |
| 纲： | 桡足纲 Copepoda |
| 目： | 剑水蚤目 Cyclopoida |
| 科： | 剑水蚤科 Cyclopidae |
| 亚科： | 剑水蚤亚科 Cyclopinae |
| 属： | 剑水蚤属 Cyclops O. F. Müller, 1785[1] |
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语源
本属的学名Cyclops源自希腊神话的独眼巨人，因为两者同样有一只大眼睛。剑水蚤属的大眼睛的颜色或红或黑。
生物学防治
Barbel及 Gambusia这两个细小鱼类的属都以剑水蚤属物种为食。在印度的卡纳塔克邦利用了这两种鱼的食性来去除可能感染了麦地那龙线虫幼虫的水蚤，从而消除透过受污染食水而引起的麦地那龙线虫病[4]。
物种
本属过往曾包括有多个亚属，但这些亚属目前均已独立出来成为各个属[1]，但在部分文献（例如：《中国动物志》）仍然可能使用旧的分类。
现时本属包括的物种计有[1]：
- Cyclops abnobensis Brehm, 1958
- Cyclops abyssicola Lilljeborg, 1901
- Cyclops abyssorum G. O. Sars, 1863
- Cyclops acanthoides Douwe, 1914
- Cyclops adolescens Herrick, 1882
- Cyclops aequoreus Fischer, 1860
- 近亲剑水蚤 Cyclops affinis G. O. Sars, 1863
- 非洲剑水蚤 Cyclops africanus Bourne, 1893
- Cyclops agilis Koch, 1838
- Cyclops agiloides G. O. Sars, 1909
- Cyclops alajensis Ulyanin, 1874
- Cyclops albicans G. W. Smith, 1909
- 白剑水蚤 Cyclops albidus Jurine, 1820
- Cyclops alius Kiefer, 1935
- Cyclops alpestris Daday, 1885
- Cyclops alticola Kiefer, 1935
- Cyclops americanus Marsh, 1893
- Cyclops amoenus Mann, 1940
- Cyclops anceps (Richard, 1897)
- Cyclops angustus S. Grassel, 1999
- Cyclops ankyrae Mann, 1940
- Cyclops anninae Menzel, 1926
- Cyclops annulatus Wierzejski, 1893
- Cyclops annulicornis Koch, 1838
- Cyclops anophthalmus Joseph, 1882
- Cyclops argulus (Fabricius, 1793)
- Cyclops armatus Tilesius, 1815
- Cyclops arnaudi G. O. Sars, 1908
- Cyclops aspericornis Daday, 1906
- Cyclops ater Herrick, 1882
- Cyclops attenuatus G. O. Sars, 1909
- Cyclops aurantius Fischer, 1860
- 澳洲剑水蚤 Cyclops australis King, 1855
- Cyclops baicalensis Vassilieva, 1950
- Cyclops bathybius Daday, 1896
- 二色剑水蚤 Cyclops bicolor G. O. Sars, 1863
- 二刺剑水蚤 Cyclops bicuspidatus Claus, 1857，二棘剑水蚤
- Cyclops bisetosus Rehberg, 1880
- Cyclops bissextilis Willey, 1925
- Cyclops bistriatus Koch, 1838
- Cyclops bodamicus Vosseler, 1886
- Cyclops bodanus Kiefer, 1954
- Cyclops bogoriensis Menzel, 1926
- Cyclops bohater Kozminski, 1933
- Cyclops bohemicus Srámek-Husek, 1938
- Cyclops bopsini Studer, 1878
- Cyclops brachypus (Kiefer, 1955)
- Cyclops bracteatus (O. F. Müller, 1776)
- Cyclops brasiliensis Dana, 1849
- Cyclops brehmi (Kiefer, 1927)
- 短尾剑水蚤 Cyclops brevicaudatus Claus, 1857
- Cyclops brevicornis Baird, 1835
- Cyclops brevicornis Claus, 1857
- Cyclops brevipes Brady, 1910
- Cyclops brevisetosus Daday, 1885
- 短棘剑水蚤 Cyclops brevispinosus Herrick, 1884
- Cyclops brucei T. Scott, 1899
- Cyclops buxtoni Gurney, 1921
- Cyclops cabanensis Russki, 1889
- Cyclops caeruleus O. F. Müller, 1776
- Cyclops canadensis Einsle, 1988
- Cyclops canthocarpoides Fischer, 1851
- Cyclops capillatus G. O. Sars, 1863
- Cyclops capilliferus S. A. Forbes, 1893
- Cyclops carolinianus Yeatman, 1944
- Cyclops caspicus Lindberg, 1942
- Cyclops castor Desmarest, 1825
- Cyclops caudatus (G. O. Sars, 1927)
- Cyclops cavernarum Ulrich, 1902
- Cyclops ceibaensis Marsh, 1919
- Cyclops cerberus Chappuis, 1934
- Cyclops charon Kiefer, 1931
- Cyclops chelifer O. F. Müller, 1776
- Cyclops chiltoni G. M. Thomson, 1883
- Cyclops christianensis Boeck, 1872
- Cyclops ciliatus G. O. Sars, 1909
- Cyclops clandestinus Yeatman, 1964
- Cyclops claudiopolitanus Daday, 1885
- Cyclops clausii Lubbock, 1863
- Cyclops clausii Poggenpol, 1874
- Cyclops claviger O. F. Müller, 1785
- Cyclops coecus Pratz, 1866
- Cyclops coeruleus O. F. Müller, 1785
- Cyclops colchidanus Borutsky, 1930
- Cyclops columbianus Grassel, 1956
- Cyclops communis Lindberg, 1938
- Cyclops compactus G. O. Sars, 1909
- Cyclops coronatus Claus, 1857
- Cyclops crassicaudis G. O. Sars, 1863
- Cyclops crassicaudoides Kiefer, 1928
- Cyclops crassicornis O. F. Müller, 1785
- Cyclops crinitus Graeter, 1910
- Cyclops croaticus Krmpotic, 1924
- Cyclops cunningtoni G. O. Sars, 1909
- Cyclops curticaudus Dana, 1847
- Cyclops curticornis O. F. Müller, 1785
- Cyclops cyprinaceus (Shaw, 1789)
- Cyclops davidi Chappuis, 1922
- Cyclops delachauxi Kiefer, 1925
- Cyclops delphinus (O. F. Müller, 1785)
- Cyclops demetiensis Scourfield, 1932
- Cyclops dengizicus Lepeshkin, 1900
- Cyclops dentatimanus Marsh, 1913
- Cyclops dentatus Rehberg, 1880
- Cyclops dentatus Koch, 1844
- Cyclops denticulatus Nicolet, 1849
- Cyclops depressus Baird, 1837
- Cyclops diaphanus Fischer, 1853
- Cyclops diminuta Lindberg, 1937
- Cyclops dimorphus Kiefer, 1934
- Cyclops distans Kiefer, 1956
- Cyclops distinctus Richard, 1887
- Cyclops diversus (Kiefer, 1935)
- Cyclops divulsus Lindberg, 1956
- Cyclops donnaldsoni Chappuis, 1929
- Cyclops dubitabilis Kiefer, 1934
- Cyclops dubius G. O. Sars, 1909
- Cyclops dulvertonensis G. W. Smith, 1909
- Cyclops dumasti Joly, 1883
- Cyclops dybowskii Landé, 1890
- Cyclops eboracensis Brady, 1902
- Cyclops echinatus Kiefer, 1926
- Cyclops ecornis Tilesius, 1819
- Cyclops edax S. A. Forbes, 1890
- Cyclops elegans Herrick, 1884
- Cyclops elgonensis Kiefer, 1932
- Cyclops elongatus Claus, 1863
- Cyclops emini Mrázek, 1898
- Cyclops entzii Daday, 1885
- Cyclops euacanthus G. O. Sars, 1909
- Cyclops ewarti Brady, 1888
- Cyclops exiguus G. O. Sars, 1909
- Cyclops exilis Coker, 1934
- Cyclops exsulis Gauthier, 1951
- Cyclops falsus Kiefer, 1929
- Cyclops fasciacornis Cragin, 1883
- Cyclops fedtschenkoi Ulyanin, 1875
- Cyclops feuerborni Kiefer, 1933
- Cyclops fimbriatus Fischer, 1853
- Cyclops finmarchicus O. F. Müller, 1776
- Cyclops fischeri Poggenpol, 1874
- Cyclops flexipes Kokubo, 1912
- Cyclops fluviatilis Herrick, 1882
- Cyclops foliaceus (Linnaeus, 1758)
- Cyclops forbesi Herrick, 1895
- Cyclops formosanus Harada, 1931
- Cyclops fragilis Kiefer, 1926
- Cyclops franciscoloi Brian, 1951
- Cyclops frivaldszkyi Daday, 1885
- Cyclops furcatus Baird, 1837
- 叉足剑水蚤 Cyclops furcifer Claus, 1857[5]
- Cyclops furi Kozhova & Pavlov, 1986
- Cyclops fuscus Jurine, 1820
- Cyclops gauthieri Green, 1963
- Cyclops gemellus (Gurney, 1928)
- Cyclops geoffroyi Samouelle, 1819
- Cyclops gibbus (Philippi, 1843)
- Cyclops gibsoni Brady, 1904
- Cyclops gigas Claus, 1857
- Cyclops glacialis Brady, 1910
- Cyclops gnatho (Philippi, 1843)
- Cyclops gracilicornis Landé, 1891
- Cyclops gracilis Lilljeborg, 1853
- Cyclops graeteri Chappuis, 1927
- Cyclops grandis Kiefer, 1935
- Cyclops grandispinifer Lindberg, 1940
- Cyclops gredleri Heller, 1871
- Cyclops gyrinus S. A. Forbes, 1890
- Cyclops hadjebensis Kiefer, 1926
- Cyclops halepensis Chappuis, 1922
- Cyclops hamatus Sovinsky, 1888
- Cyclops harpacticoides Shmankevich, 1875
- Cyclops haueri Kiefer, 1931
- Cyclops heberti Einsle, 1996
- Cyclops helgolandicus Rehberg, 1880
- Cyclops helleri Brady, 1878
- Cyclops horvathii Daday, 1885
- Cyclops hungaricus Daday, 1885
- Cyclops hutchinsoni Kiefer, 1936
- Cyclops hypogeus Kiefer, 1930
- Cyclops ignaeus Poggenpol, 1874
- Cyclops igneus Poggenpol, 1874
- Cyclops incertus Wolf, 1905
- Cyclops indolusitanicus Lindberg, 1938
- Cyclops inermis Tilesius, 1812
- Cyclops infernus Kiefer, 1930
- Cyclops ingens Herrick, 1882
- Cyclops inopinatus (G. O. Sars, 1927)
- Cyclops inopinus Kiefer, 1926
- Cyclops insectus S. A. Forbes, 1882
- Cyclops insignis Claus, 1857
- Cyclops intermedius Sovinsky, 1888
- Cyclops isodactylus (Philippi, 1843)
- Cyclops jashnovi Streletskaya, 1990
- Cyclops javanus Kiefer, 1930
- Cyclops jeanneli Chappuis, 1929
- Cyclops johnstoni Baird, 1834
- Cyclops josephi Moniez, 1887
- Cyclops juri Parveen, Mahoon & Saleem, 1990
- Cyclops karamani Kiefer, 1932
- Cyclops karvei Kiefer & Moorthy, 1935
- Cyclops kaufmanni Ulyanin, 1875
- Cyclops kentanensis Harada, 1931
- Cyclops kieferi Chappuis, 1925
- Cyclops kievensis Sovinsky, 1887
- Cyclops kikuchii Smirnov, 1932
- Cyclops kolensis Lilljeborg, 1901
- Cyclops korostyschevi Sovinsky, 1888
- Cyclops kozminskii Lindberg, 1942
- Cyclops krillei Studer, 1878
- Cyclops lacinulatus O. F. Müller, 1776
- Cyclops lacunae Lowndes, 1926
- Cyclops lacustris G. O. Sars, 1863
- 拉达克剑水蚤 Cyclops ladakanus Kiefer, 1936
- Cyclops laevimargo G. O. Sars, 1909
- Cyclops landei Mahoon & Zia, 1985
- Cyclops languidoides Lilljeborg, 1901
- Cyclops languidulus Willey, 1925
- Cyclops languidus (G. O. Sars, 1863)
- Cyclops lascivus Poggenpol, 1874
- Cyclops laticauda Templeton, 1836
- Cyclops latipes Lowndes, 1927
- Cyclops latissimus Poggenpol, 1874
- Cyclops laurenticus Lindberg, 1956
- Cyclops learii Ulrich, 1902
- Cyclops leewenhoekii Hoek, 1878
- Cyclops leptopus Kiefer, 1927
- Cyclops leuckarti Claus, 1857
- Cyclops lilljeborgi G. O. Sars, 1918
- Cyclops linjanticus Kiefer, 1928
- Cyclops littoralis Brady, 1872
- Cyclops lobulosus Ekman, 1905
- Cyclops longicaudatus Poggenpol, 1874
- Cyclops longispina Templeton, 1836
- Cyclops longistylis Brady, 1907
- Cyclops lubbocki Brady, 1869
- Cyclops lucidulus Koch, 1838
- Cyclops maarensis Vosseler, 1889
- Cyclops macleayi Dana, 1847
- Cyclops macruroides Lilljeborg, 1901
- Cyclops macrurus G. O. Sars, 1863
- Cyclops macuroides Lilljeborg, 1901
- Cyclops madagascariensis Kiefer, 1926
- Cyclops magniceps Lilljeborg, 1853
- Cyclops magnoctavus Cragin, 1883
- 大剑水蚤 Cyclops magnus (Marsh, 1920)
- Cyclops malayicus Kiefer, 1930
- Cyclops margaretae Lindberg, 1938
- Cyclops margoi Daday, 1885
- Cyclops marinus Prestandrea, 1833
- Cyclops matritensis Velasquez, 1941
- Cyclops mendocinus Wierzejski, 1893
- Cyclops menzeli Kiefer, 1926
- Cyclops meridianus Kiefer, 1926
- Cyclops michaelseni Mrázek, 1901
- Cyclops micropus Kiefer, 1932
- Cyclops miles Nicolet, 1849
- Cyclops miniatus Lilljeborg, 1901
- Cyclops minimus Kiefer, 1933
- Cyclops minnilus S. A. Forbes, 1893
- Cyclops minuticornis O. F. Müller, 1785
- Cyclops minutissimus Kiefer, 1933
- Cyclops minutus Claus, 1863
- Cyclops minutus O. F. Müller, 1776
- Cyclops modestus Herrick, 1883
- Cyclops moghulensis Lindberg, 1939
- Cyclops monacanthus Kiefer, 1928
- Cyclops monardi Perret, 1925
- Cyclops mongoliensis Einsle, 1992
- Cyclops mulleri Ferussac, 1806
- Cyclops muscicola Menzel, 1926
- Cyclops muscicolus Lastochkin, 1927
- Cyclops nanus G. O. Sars, 1863
- Cyclops naviculus Say, 1818
- Cyclops navus Herrick, 1882
- Cyclops nearcticus Kiefer, 1934
- Cyclops necessarius Kiefer, 1926
- Cyclops neglectus G. O. Sars, 1909
- Cyclops neumani Pesta, 1927
- Cyclops neymanae Streletskaya, 1990
- Cyclops niceae Mann, 1940
- Cyclops nigeriae Brady, 1910
- Cyclops nigricauda Norman, 1869
- Cyclops nivalis Daday, 1885
- Cyclops novaezealandiae G. M. Thomson, 1879
- Cyclops nubicus Chappuis, 1922
- Cyclops obesicornis Templeton, 1836
- Cyclops obsoletus Koch, 1838
- Cyclops ochridanus Kiefer, 1932
- Cyclops odessanus Shmankevich, 1875
- 长刺剑水蚤 Cyclops oithonoides G. O. Sars, 1863
- Cyclops oligarthrus G. O. Sars, 1909
- Cyclops operculatus Chappuis, 1917
- 东方剑水蚤 Cyclops orientalis Ulyanin, 1875
- Cyclops ornatus Poggenpol, 1874
- Cyclops ovalis Brady, 1872
- Cyclops pachycomus G. O. Sars, 1909
- Cyclops pallidus Norman, 1869
- Cyclops paludicola Herbst, 1959
- Cyclops palustris Sovinsky, 1888
- Cyclops panamensis Marsh, 1913
- Cyclops papuanus Daday, 1901
- Cyclops paradyi Daday, 1885
- Cyclops paraplesius Kiefer, 1929
- Cyclops parcus Herrick, 1882
- Cyclops pauper Fric, 1871
- Cyclops pectinatus Herrick, 1883
- Cyclops pectinifer Cragin, 1883
- Cyclops pelagicus (Rose, 1929)
- Cyclops pennatus Claus, 1857
- Cyclops pentagonus Vosseler, 1886
- Cyclops perarmatus Cragin, 1883
- Cyclops phaleratus Koch, 1838
- Cyclops phaleroides Labbé, 1927
- Cyclops philippinensis Marsh, 1932
- Cyclops phreaticus Chappuis, 1928
- Cyclops pictus Koch, 1838
- Cyclops pilosus Kiefer, 1934
- Cyclops piscinus (Linnaeus, 1761)
- Cyclops planus Gurney, 1909
- Cyclops plumosus (Philippi, 1843)
- Cyclops poggenpolii Sovinsky, 1888
- Cyclops poppei Rehberg, 1880
- Cyclops potamius Burckhardt, 1913
- 葱绿剑水蚤 Cyclops prasinus Fischer, 1860
- Cyclops prealpinus Kiefer, 1939
- Cyclops procerus Herbst, 1955
- Cyclops productus (O. F. Müller, 1785)
- Cyclops prolatus Kiefer, 1935
- Cyclops pubescens Dana, 1847
- Cyclops purpureus (Philippi, 1843)
- Cyclops pusillus Brady, 1904
- Cyclops puteanus Frey, 1869
- Cyclops pygmaeus Rehberg, 1880
- Cyclops quadricornis (Linnaeus, 1758)
- Cyclops quinquepartitus Marsh, 1913
- Cyclops racovitzai Chappuis, 1923
- Cyclops rarispinus G. O. Sars, 1909
- Cyclops reductus Chappuis, 1925
- Cyclops restrictus Lindberg, 1948
- Cyclops ricae Monchenko, 1977
- Cyclops richardi Lindberg, 1942
- Cyclops robustus G. O. Sars, 1863
- Cyclops roseus Daday, 1885
- Cyclops rostratus (Philippi, 1843)
- Cyclops roumaniae Cosmovici, 1900
- Cyclops royi Lindberg, 1940
- Cyclops rubellus Lilljeborg, 1901
- Cyclops rubens O. F. Müller, 1785
- Cyclops salinus Brady, 1903
- Cyclops salmoneus (J. C. Fabricius, 1792)
- Cyclops saltatonius (O. F. Müller, 1776)
- Cyclops sanfilippoi Brian, 1951
- Cyclops sanguineus (Philippi, 1843)
- Cyclops schmeili Poppe & Mrázek, 1895
- Cyclops scourfieldi Brady, 1891
- Cyclops scutifer G. O. Sars, 1863
- Cyclops semiserratus G. O. Sars, 1909
- Cyclops sensitivus Graeter & Chappuis, 1914
- Cyclops serratus Pratz, 1866
- Cyclops serrulatoides Labbé, 1927
- Cyclops serrulatus Fischer, 1851
- Cyclops setiger Frey, 1869
- Cyclops setosus Haldeman, 1842
- Cyclops sevani Meshkova, 1947
- Cyclops shatalovi Streletskaya, 1990
- 标志剑水蚤 Cyclops signatus Koch, 1838
- Cyclops silesicus Schäfer, 1934
- Cyclops silvestrii Brian, 1927
- 长腹剑水蚤 Cyclops similis Templeton, 1836
- Cyclops simillimus Brady, 1907
- 简剑水蚤 Cyclops simplex Poggenpol, 1874
- Cyclops skopljensis Kiefer, 1932
- Cyclops smirnovi Rylov, 1948
- Cyclops soli Kokubo, 1912
- Cyclops spartinus Ruber, 1966
- 分离剑水蚤 Cyclops speratus Lilljeborg, 1901
- Cyclops spinifer Daday, 1902
- Cyclops spinulosus Claus, 1893
- Cyclops stagnalis Einsle, 1996
- Cyclops staheli Chappuis, 1917
- Cyclops staphylinus Desmarest, 1825
- 英勇剑水蚤 Cyclops strenuus Fischer, 1851
- Cyclops stroemii Baird, 1834
- Cyclops stuhlmanni Mrázek, 1895
- Cyclops stygius Chappuis, 1924
- Cyclops subaequalis Kiefer, 1928
- Cyclops subterraneus Pratz, 1866
- Cyclops subtropicus Lindberg, 1937
- Cyclops sumatranus Kiefer, 1933
- Cyclops sydneyensis Schmeil, 1898
- Cyclops sylvestrii Brian, 1927
- Cyclops taipehensis Harada, 1930
- Cyclops tanganicae (Gurney, 1928)
- Cyclops tatricus Kozminski, 1927
- Cyclops tenellus G. O. Sars, 1909
- Cyclops tenuicaudis Daday, 1885
- Cyclops tenuicornis Claus, 1857
- Cyclops tenuipes (Philippi, 1843)
- Cyclops tenuis Marsh, 1910
- Cyclops tenuisaccus G. O. Sars, 1927
- Cyclops tenuissimus Herrick, 1883
- Cyclops teras Graeter, 1907
- Cyclops thomasi S. A. Forbes, 1882
- Cyclops thorace (Ström, 1770)
- Cyclops transilvanicus Daday, 1885
- Cyclops tricolor Lindberg, 1937
- Cyclops trisetosus Herbst, 1957
- Cyclops triumvirorum Kiefer, 1935
- Cyclops troglodytes Chappuis, 1923
- Cyclops troglophilus Kiefer, 1932
- Cyclops tropicus Kiefer, 1932
- Cyclops trouchanowi Sovinsky, 1888
- Cyclops uljanini Sovinsky, 1887
- Cyclops uniangulatus Cragin, 1883
- Cyclops unisetiger Graeter, 1899
- Cyclops uruguayensis Kiefer, 1935
- Cyclops varicans G. O. Sars, 1863
- Cyclops varicoides Brady, 1908
- 变色剑水蚤 Cyclops varius Lilljeborg, 1901
- Cyclops venustoides Coker, 1934
- Cyclops venustus Norman & T. Scott, 1906
- 春季剑水蚤 Cyclops vernalis Fischer, 1853
- 近邻剑水蚤 Cyclops vicinus Ulyanin, 1875
- 近邻剑水蚤 Cyclops vicinus vicinus
- 叶片剑水蚤 Cyclops vicinus lobosus
- Cyclops viduus Kiefer, 1933
- 文森特剑水蚤 Cyclops vincentianus Brian, 1927
- Cyclops vinceus Shmankevich, 1875
- Cyclops virescens Brady, 1910
- 绿剑水蚤 Cyclops viridis (Jurine, 1820)
- Cyclops viridosignatus E. F. Byrnes, 1909
- Cyclops vitiensis Dana, 1847
- Cyclops vulgaris Koch, 1838
- Cyclops wigrensis Streletskaya, 1988
- Cyclops zschokkei Graeter, 1910
流行文化
参考文献
- ^ 1.0 1.1 1.2 1.3 Daphne Cuvelier & T. Chad Walter. Cyclops Müller, 1785. World Copepoda database.
- ^ G. G. Marten. Issues in the development of Cyclops for mosquito control. M. F. Uren, J. Blok & L.H. Manderson (编). Arbovirus Research in Australia: Proceedings of the Fifth Symposium (August 28 - September 1, 1989, Brisbane, Australia) (PDF). 1986: 159–164 [2019-10-03]. （原始内容存档 (PDF)于2020-01-14）.
- ^ 無節蚴；無節幼蟲 nauplius. 台湾国家教育研究院双语词汇、学术名词暨辞书资讯网. [2020-07-27] （中文（繁体））.
- ^ USA National Research Council; World Health Organization. Workshop on opportunities for control of dracunculiasis, 16-19 June 1982, Washington, D.C. Contributed papers (PDF). Washington, D.C.: National Academy Press. 1985: 153–177 [2019-10-03]. （原始内容存档 (PDF)于2017-03-12）.
- ^ 中国科学院动物研究所. 叉足剑水蚤. 《中国动物物种编目数据库》. 中国科学院微生物研究所. [2009-04-28]. （原始内容存档于2016-03-05） （中文（简体））.
